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Glossary

Botanical, ecological and geological terms used on this site, explained.

A

Achene Botany

A small, dry, one-seeded fruit that does not split open at maturity.

Aeolian Landscape & ecology

Wind-driven. Used to describe landforms, sediments and geological processes shaped by the action of wind.

Alien species Botany

A species introduced, deliberately or accidentally, by people outside its natural range. Also called non-native or exotic species.

Alluvial fan Landscape & ecology

A low, outspread, relatively flat to gently sloping mass of loose rock material, shaped like a fan or a segment of a cone, deposited by streams at the mouths of tributary valleys onto a plain or broad valley.

Alternate Botany

Arranged singly at different heights along a stem, rather than in pairs.

Amplexicaul Botany

Having a leaf base that clasps or partly surrounds the stem.

Annual Botany

A plant that completes its whole life cycle, from seed to flowering, fruiting and death, within one year. In AlUla, many annuals germinate only after rain.

Ascending Botany

Growing upward from a base that initially lies or spreads along the ground.

B

Biological soil crust Botany

A thin living layer at the soil surface, made of cyanobacteria, algae, lichens and mosses, that binds soil particles, reduces erosion and enriches the soil.

Bioregion Landscape & ecology

A geographical area defined by characteristics of the natural environment. Saudi Arabian bioregions (Llewellyn, 2013) are based on physiography (geology and topography), altitude, climate, soils, and vegetation structure and composition.

Biotype Landscape & ecology

Similar to a habitat but defined in relation to a suite of species.

Bract Botany

A modified leaf associated with a flower or flower cluster.

Bryophyte Botany

A small, non-flowering land plant without true roots or vascular tissue: mosses, liverworts and hornworts. In AlUla, 48 bryophyte species are recorded, mostly in shaded and moist refuges.

Butte Landscape & ecology

An isolated hill with steep sides and a flat, slightly rounded top.

C

Caducous Botany

Falling off early or before the end of the growing season.

Calyx Botany

The outermost flower whorl, usually made up of sepals.

Canescent Botany

Covered with a fine, often grayish-white covering of hairs.

Canyon Landscape & ecology

A large chasm between cliffs.

Capitulum Botany

A compact head of many small flowers, as in daisies.

Capsule Botany

A dry fruit that splits open at maturity to release its seeds.

Chamaephyte Botany

A low shrub or dwarf shrub whose regrowth buds sit close to the ground (below about 25 cm), where they are sheltered from wind and heat. Many desert subshrubs are chamaephytes.

Clasts Landscape & ecology

Fragments or grains of pre-existing rocks and minerals.

Climber Botany

A plant that climbs on rocks or other plants for support, using twining stems, tendrils or hooks.

Cordate Botany

Heart-shaped, usually with a notch at the base.

Coriaceous Botany

Leathery in texture.

Corolla Botany

The flower’s petals, considered together.

Crater Landscape & ecology

The top of a volcano, which is lower than the surrounding rim.

Cryptophyte Botany

A plant whose regrowth buds are hidden below the soil surface or under water during the unfavourable season. Geophytes are the land-dwelling cryptophytes.

Cyme Botany

A flower cluster in which the central or terminal flower opens first.

D

Data Deficient Botany

IUCN Red List category for a species whose risk of extinction cannot be assessed because too little is known about its distribution or populations. Many plants of AlUla are in this situation.

Decreaser species Landscape & ecology

Species present in areas with low to no degradation, balanced ecosystem dynamics and controlled grazing. They are often palatable, and are called "decreasers" because they tend to decrease as environmental degradation intensifies.

Degradation Landscape & ecology

The process by which the quality of a natural habitat is reduced, making it less able to support wildlife. It can have human or natural causes; on this site, it refers to human causes.

Dehiscent Botany

Opening naturally at maturity to release seeds.

E

Ecosystem type Landscape & ecology

A complex of organisms and their associated physical environment, united by similar ecological processes, whose biotic structure and composition differ substantially from those of other ecosystem types.

Endemic species Landscape & ecology

A plant or animal found naturally only in a specific geographic region. For example, national endemics are found only in Saudi Arabia, and Arabian endemics only in the countries of the Arabian Peninsula.

Ephemeral Botany

Lasting for a very short time. In plants, it often describes species that grow, flower, and set seed quickly when conditions are favorable.

Erect Botany

Growing upright.

F

Family Botany

A group of related genera sharing common features and ancestry. Plant family names end in "-aceae", for example Fabaceae (legumes) or Asteraceae (daisies).

Flowering Botany

The period during which a plant bears open flowers. In AlUla, most plants flower in late winter and spring, after the winter rains.

Follicle Botany

A dry fruit that opens along one side to release its seeds.

Fruiting Botany

The period during which a plant bears fruits and seeds. Also called fructification.

G

Genus Botany

A group of closely related species (plural: genera). The genus is the first part of a scientific name, for example Vachellia in Vachellia tortilis.

Geographically restricted Landscape & ecology

Fauna species with a limited distribution, defined as an Area of Occupancy (AOO) under 10 km² and/or an Extent of Occurrence (EOO) under 100 km², following the IUCN methodology.

Geomorphology Landscape & ecology

The branch of geology concerned with the structure, origin and development of the topographical features of the earth's surface.

Geophyte Botany

A plant that survives unfavorable seasons using underground storage organs, such as bulbs or tubers.

Glabrous Botany

Smooth and without hairs.

Glandular Botany

Bearing glands that may produce or release sticky or aromatic substances.

Growth form Botany

The overall shape and structure of a plant (tree, shrub, herb, climber…), and the way it survives the unfavourable season. On this site, growth forms follow the Raunkiær life-form system: see Phanerophyte, Chamaephyte, Hemicryptophyte, Cryptophyte, Geophyte and Therophyte.

H

Habitat Landscape & ecology

Plant and animal communities, as the characterising elements of the biotic environment, together with abiotic factors (soil, climate, water availability and quality, and others), operating together at a particular scale (EUNIS definition, European Environment Agency, 2023).

Halophyte Botany

A plant adapted to grow in salty soils, such as those of ephemeral flooded depressions.

Harmonised Landscape & ecology

Describes data aligned and integrated from different sources, methodologies or classification systems to create a consistent and comparable dataset.

Harrat Landscape & ecology

A lava field forming a high plateau. Harrat surfaces clearly reflect their age: older harrats are severely weathered and eroded, while younger ones show little erosion and can appear as an impassable, chaotic jumble of angular blocks.

Hemicryptophyte Botany

A perennial plant whose above-ground parts die back in the unfavourable season, leaving regrowth buds at the soil surface, often protected by dead leaves. Many perennial grasses are hemicryptophytes.

Herbaceous plant Botany

A plant with soft, non-woody stems. Herbaceous plants can be annual, ephemeral or perennial.

Hispid Botany

Covered with stiff, bristly hairs.

I

IUCN Red List Botany

The global inventory of the conservation status of species, maintained by the International Union for Conservation of Nature (IUCN). Its categories, from least to most at risk, are: Least Concern, Near Threatened, Vulnerable, Endangered, Critically Endangered, Extinct in the Wild and Extinct. Species not yet assessed are Not Evaluated; those lacking data are Data Deficient.

Increaser species Landscape & ecology

Species, possibly native to Saudi Arabia (so not alien invasives), that spread quickly with environmental degradation or overgrazing. Some are unpalatable to grazers, and some compete with established plant communities as degradation begins or intensifies.

Inflorescence Botany

The arrangement or cluster of flowers on a plant.

Inselberg Landscape & ecology

An isolated rock hill, ridge or small mountain that rises abruptly from a virtually level surrounding plain.

Invasive species Botany

An alien species that spreads rapidly in its new range and harms native species, habitats or human activities.

L

Lamellar Landscape & ecology

Describes rocks, minerals or soils composed of fine, alternating, parallel layers or flat plates.

Lanceolate Botany

Shaped like a lance: longer than wide and tapering toward the ends.

Lava flow Landscape & ecology

A stream or sheet of molten or solidified lava erupted from a volcano onto the Earth's surface. Solidified lava flows can be described as fresh or eroded.

Lichen Botany

A partnership between a fungus and an alga or cyanobacterium, forming a crust, scales or small leafy or shrubby growths on rocks and soil.

M

Mesa Landscape & ecology

A flat-topped hill with steep sides, smaller than a plateau and with much steeper sides. In AlUla, mesas are typically in sedimentary rocks, while plateaus are the harrats.

Mountain Landscape & ecology

A naturally raised landform with steep, sharp edges and a peak.

Mucronate Botany

Ending in a short, abrupt, sharp point.

N

Nabkha Landscape & ecology

An accumulation of wind-blown sediment around a plant, forming a raised, fertile mound.

Native species Botany

A species that occurs naturally in a region, without having been introduced by people.

Nutlet Botany

A small, usually hard, one-seeded fruit or fruit section.

O

Obovate Botany

Egg-shaped with the broader end toward the tip.

Opposite Botany

Arranged in pairs directly across from one another on a stem.

Outcrop Landscape & ecology

An area of exposed rock sticking out from the surrounding ground.

Ovary Botany

The part of a flower that contains ovules and can develop into fruit after fertilization.

Overgrazing Landscape & ecology

Intensive grazing by animals on an area of land, particularly rangelands and grasslands, to the point that vegetation is damaged and the soil becomes liable to erosion by wind and rain, sometimes resulting in desertification.

P

Palatable Botany

Describes plants that livestock and wildlife readily eat. Palatable species tend to decline under heavy grazing.

Panicle Botany

A branched inflorescence in which the flower stalks are themselves branched.

Pappus Botany

A tuft or ring of hairs or bristles on a fruit, often helping it disperse by wind.

Parasite Botany

A plant that draws all or part of its water and nutrients from another living plant (its host), through specialised organs attached to the host's roots or stems.

Pedicel Botany

The small stalk that supports an individual flower within a cluster.

Pediment Landscape & ecology

A gently sloping bedrock surface that forms at the base of a mountain or hill, often covered by a thin layer of gravel or alluvium.

Pediplain Landscape & ecology

An extensive, gently sloping plain formed by the joining of multiple pediments at the base of a mountain range.

Pedogenesis Landscape & ecology

The process of soil formation.

Perennial Botany

A plant that lives for more than two years.

Perianth Botany

The flower’s non-reproductive parts; it includes the sepals and petals, or similar structures when these are not distinct.

Petiolate Botany

Having a petiole, the stalk that attaches a leaf blade to a stem.

Phanerophyte Botany

A woody plant, tree or tall shrub, whose regrowth buds are carried well above the ground (more than about 25 cm). Example: acacias (Vachellia).

Phenology Botany

The timing of recurring stages in a plant's life, such as flowering and fruiting, through the year. On species pages, the phenology chart shows in which months each species has been observed in flower or in fruit in AlUla.

Pinnatisect Botany

Deeply divided into segments arranged along both sides of a central axis.

Plateau Landscape & ecology

A large flat expanse that rises, often steeply, above the surrounding landscape.

Prostrate Botany

Growing flat or close to the ground.

Pubescent Botany

Covered with short, soft hairs.

Pyroduct Landscape & ecology

A natural tube formed by flowing lava. When emptied of lava it can resemble an elongated cave. Also called a lava tube.

Q

Quweria Sandstone Landscape & ecology

A sandstone formation in AlUla with honey-coloured, fairly massive layers, dominated by fluvial deposits and quartz sediment. Its cement is relatively weakened by hydrolysis and chemical weathering.

R

Raceme Botany

An elongated flower cluster with stalked flowers arranged along a central stem.

Range-restricted Landscape & ecology

Refers to species known to occur only in Saudi Arabia and neighbouring countries. They are not regional endemics but have a restricted range.

Raunkiær life forms Botany

A classification of plants devised by the Danish botanist Christen Raunkiær, based on where the buds that allow regrowth are placed during the unfavourable season (drought or cold): high above the ground, near the ground, at the soil surface, below ground, or only in seeds.

Refugia Landscape & ecology

Areas to which components of biodiversity retreat, in which they persist, and from which they can potentially expand under changing environmental conditions.

Rhizome Botany

A horizontal underground stem that can produce roots and new shoots.

Runnel Landscape & ecology

A small stream or channel.

S

Sand sheet Landscape & ecology

A low-relief sand surface largely devoid of dunes, although it may show low-relief aeolian features.

Saq Sandstone Landscape & ecology

The youngest sandstone formation in AlUla: medium-grained, well-rounded sands with graded bedding, interlaid with shallow marine deposits. Its cement is relatively weakened by hydrolysis and chemical weathering, while the marine deposits form hard iron beds.

Scabrid Botany

Rough to the touch because of short, stiff hairs or projections.

Schizocarp Botany

A dry fruit that splits into separate one-seeded sections when mature.

Scientific name Botany

The universal Latin name of a taxon, written in italics and followed by the name of the author(s) who first described it, for example Morettia parviflora Boiss.

Serrate Botany

Having saw-like teeth along the edge, typically pointing toward the tip.

Sessile Botany

Attached directly to a stem, without a stalk.

Shear zone Landscape & ecology

A zone where strain forces opposing sides of the rock to slide past each other, here as a result of faulting linked to the opening of the Red Sea. The rock there is broken and more brittle, and erosion works faster.

Shrub Botany

A woody perennial plant, smaller than a tree, usually with several stems branching from the base. Dwarf shrubs and subshrubs are low shrubs, often woody only at the base.

Siq Sandstone Landscape & ecology

The oldest sandstone formation in AlUla, made of several thick, medium to coarse-grained beds with large cross-bedding throughout, indicating high-energy river channels.

Species Botany

The basic unit of classification: a group of individuals that resemble each other and can interbreed. A species is named with two words, the genus and the species epithet, for example Rhazya stricta.

Stamen Botany

The pollen-producing part of a flower, usually consisting of a filament and anther.

Stratum Botany

The vertical layer of vegetation a plant belongs to: trees, shrubs, herbaceous plants or climbers. On this site, the stratum is used as a simple filter in the species list.

Subshrub Botany

A small shrub with woody growth mainly at the base and softer stems above.

Subspecies Botany

A subdivision of a species, usually a population with distinct features and its own geographic range (abbreviated subsp.). A variety (var.) is a lower rank still.

Substrate Landscape & ecology

In ecology, the substance, surface or base on which biological processes occur or an organism lives.

Succulent Botany

Having fleshy tissues adapted to store water.

Synonym Botany

Another scientific name that has been used for the same taxon but is no longer the accepted one.

T

Tafoni Landscape & ecology

Rounded, smooth cavities or hollows in rock, often grouped and described as honeycomb weathering.

Taxon Botany

Any unit of biological classification, such as a family, genus, species or subspecies (plural: taxa).

Therophyte Botany

A plant that survives the unfavourable season only as a seed: an annual or ephemeral. Therophytes make up a large part of desert floras.

Threatened species Botany

In the IUCN Red List, a species classified as Vulnerable, Endangered or Critically Endangered.

Tomentose Botany

Densely covered with soft, woolly hairs.

Tree Botany

A woody perennial plant, usually with a single main trunk, that reaches several metres in height.

Tuber Botany

A swollen underground stem or root that stores nutrients and may help the plant survive or reproduce.

U

Umbel Botany

A flower cluster in which stalks arise from one point, spreading like the ribs of an umbrella.

Un-weathered lava flow Landscape & ecology

A fresh lava flow, where the basalt has not been significantly eroded.

V

Vegetation type Landscape & ecology

The dominant plant species and their structural characteristics in an area.

Volcanic cone Landscape & ecology

A hill-shaped landform that forms around a volcano.

W

Wadi Landscape & ecology

A river channel, which can also include the floodplain and valley of a stream, usually dry except during the rainy season.

X

Xerophyte Botany

A plant adapted to survive with very little water, for example through small or absent leaves, thick cuticles, deep roots or water-storing tissues.